Skip to content Skip to navigation


Bojian Zhong Professor

Research Interests: 
  • Plant biodiversity and adaptive evolution
  • Plant phylogenomics
  • Evolutionary genomics

Educational Background

PhD, Plant Evolution, Massey University (2014)

MS, Botany, Fudan University (2010)

BS, Life Sciences, China University of Geosciences (2007)

Research Projects

  • The National Natural Science Foundation of China, 2020-2023
  • The National Natural Science Foundation of China, 2016-2019
  • The Key Programme of Science Research Project of Jiangsu Province, 2016-2019
  • Six Talent Peaks Project of Jiangsu Province, 2016-2019
  • The Natural Science Foundation of Jiangsu Province, 2015-2018
  • The Priority Academic Program Development of Jiangsu Higher Education Institutions, 2014-2017

Journal Articles

1.    Zhang Z†, Qu C†, Zhang K†, He Y†, Zhao X†, Yang L†, … , Leliaert F, Bhattacharya D, De Clerck O, Zhong B* (Lead Contact), Miao J* (2020). Adaptation to extreme Antarctic environments revealed by the genome of a sea ice green alga. Current Biology. 30: 3330-3341.
2.    Nie Y, Foster C, Zhu T, Yao R, Duchêne D, Ho S, Zhong B* (2020). Accounting for uncertainty in the evolutionary timescale of green plants through clock-partitioning and fossil calibration strategies. Systematic Biology. 69: 1-16. 
3.    Yang L, Su D, Chang X, Foster C, Sun L, Huang C, Zhou X, Zeng L, Ma H, Zhong B* (2020). Phylogenomic insights into deep phylogeny of angiosperms based on broad nuclear gene sampling. Plant Communications. 1: 100027.
4.    Zhang Z, Chang X, Su D, Yao R, Liu X, Zhu H, Liu G, Zhong B* (2020). Comprehensive transcriptome analyses of two Oocystis algae provide insights into the adaptation to Qinghai-Tibet Plateau. Journal of Systematics and Evolution.
5.    Han X, Chang X, Zhang Z, Chen H, He H*, Zhong B*, Deng X* (2019). Origin and evolution of core components responsible for monitoring light environment changes during plant terrestrialization. Molecular Plant. 12: 847-862. 
6.    Zhang Z†, Qu C†, Yao R, Nie Y, Xu C, Miao J, Zhong B* (2019). The parallel molecular adaptations to the Antarctic cold environment in two psychrophilic green algae. Genome Biology and Evolution. 11: 1897-1908. 
7.    Fang L, Leliaert F, Novis P, Zhang Z, Zhu H, Liu G, Penny D, Zhong B* (2018). Improving phylogenetic inference of core Chlorophyta using chloroplast sequences with strong phylogenetic signals and heterogeneous models. Molecular Phylogenetics and Evolution. 127: 248-255. 
8.    Zhang Z, An M, Miao J, Gu Z, Liu C, Zhong B* (2018). The Antarctic sea ice alga Chlamydomonas sp. ICE-L provides insights into adaptive patterns of chloroplast evolution. BMC Plant Biology. 18: 53. 
9.    Zhong, B*., Betancur-R, R*. (2017). Expanded taxonomic sampling coupled with gene genealogy interrogation provides unambiguous resolution for the evolutionary root of angiosperms. Genome Biology and Evolution. 9: 3154-3161. 
10.    Fang, L., Leliaert, F., Zhang, Z., Penny, D., Zhong, B*. (2017). Evolution of the Chlorophyta: Insights from chloroplast phylogenomic analyses. Journal of Systematics and Evolution. 55: 322-332.
11.    Sun, L., Fang, L., Zhang, Z., Penny, D., Zhong, B*. (2016). Chloroplast Phylogenomic Inference of Green Algae Relationships. Scientific Reports. 6: 20528.
12.    Finet, C*., Floyd, S.K., Conway, S.J., Zhong, B., Scutt, C.P., Bowman, J.L. (2016) Evolution of the YABBY gene family in seed plants. Evolution & Development. 18: 116-126.
13.    Edwards, S.V*., Xi, Z., Janke, A., Faircloth, B.C., McCormack, J.E., Glenn, T.C., Zhong, B., Wu, S., Lemmon, E.M., Lemmon, A.R., Leaché, A.D., Liu, L., Davis, C.C. (2016). Implementing and testing the multispecies coalescent model: A valuable paradigm for phylogenomics. Molecular Phylogenetics and Evolution. 94: 447-462.
14.    Zhong, B*., Sun, L., and Penny, D. (2015). The origin of land plants: a phylogenomic perspective. Evolutionary Bioinformatics. 11: 137-141.
15.    Penny, D*., Zhong, B. (2015). Two fundamental questions about protein evolution. Biochimie. 119: 278-283.
16.    Zhong, B*., Liu, L*., and Penny, D. (2014). The multispecies coalescent model and land plant origins: a reply to Springer and Gatesy. Trends in Plant Science. 19: 270-272.
17.    Zhong, B*., Xi, Z., Goremykin, V.V., Fong, R., McLenachan, T., Novis P.M., Davis C.C., and Penny, D. (2014). Streptophyte algae and the origin of land plants revisited using heterogeneous models with three new algal chloroplast genomes. Molecular Biology and Evolution. 31: 177-183.
18.    Zhong, B*., Fong, R., Collins L.J., McLenachan, T., and Penny, D. (2014). Two new fern chloroplasts and decelerated evolution linked to the long generation time in tree ferns. Genome Biology and Evolution. 6: 1166-1173.
19.    Zhong, B*., Liu, L., Yan, Z., and Penny, D. (2013). Origin of land plants using the multispecies coalescent model. Trends in Plant Science. 18: 492-495.
20.    Goremykin,V.V*., Nikiforova, S.V., Biggs, P.J. Zhong, B., DeLange, P., Martin,W., Woetzel, S., Atherton, R.A., McLenachan, T., Lockhart P.J. (2013). The evolutionary root of flowering plants. Systematic Biology. 62: 51-62.
21.    White, W.T.J†., Zhong, B†., Penny, D*. (2013). Beyond reasonable doubt: evolution from DNA sequences. PLoS ONE. 8: e69924. (+equal contribution)
22.    Zhong, B*., Deusch, O., Goremykin, V.V., Penny, D., Biggs, P.J., Atherton, R.A., Nikiforova, S.V., Lockhart, P.J. (2011). Systematic error in seed plant phylogenomics. Genome Biology and Evolution. 3:1340-1348.
23.    Zhong, B*., Yonezawa, T., Zhong, Y., Hasegawa, M*. (2010). The position of Gnetales among seed plants: Overcoming pitfalls of chloroplast phylogenomics. Molecular Biology and Evolution. 27: 2855-2863.
24.    Zhong, B., Yonezawa, T., Zhong, Y., Hasegawa, M*. (2009). Episodic evolution and adaptation of chloroplast genomes in ancestral grasses. PLoS ONE. 4: e5297.
25.    Hasegawa, M*., Zhong, B., Zhong, Y. (2009). Adaptive evolution of chloroplast genomes in ancestral grasses. Plant Signaling & Behavior. 4: 623-624.

Honors and Awards

Top Academic Leaders of Qing Lan Projects of Jiangsu, 2016

Hundred Talent Program of Nanjing Normal University, 2014

Dean’s List of Exceptional Doctoral Theses (Top 5%), 2014

Chinese Government Award for Outstanding Self-financed Students Abroad, 2012

Outstanding Master Thesis Award of Fudan University, 2011

Academic Title

  1. Associate Editor, Journal of Molecular Evolution
  2. Editor, Journal of Systematics and Evolution